Classification
 Nomenclature
Scientific Name:
Veronica raoulii Hook.f., Handb. New Zealand Fl. 214 (1864)
Synonymy:
  • Hebe raoulii (Hook.f.) Cockayne & Allan, Trans. New Zealand Inst. 57: 44 (1926)
  • Heliohebe raoulii (Hook.f.) Garn.-Jones, New Zealand J. Bot. 31: 333 (1993)
  • Parahebe raoulii (Hook.f.) Heads, Bot. J. Linn. Soc. 115: 81 (1994)
Lectotype (designated by Garnock-Jones 1993): Akaroa, New Zealand, Raoul. The specimen so labelled was probably collected in Canterbury by Dr Lyall, K
Etymology:
Named after Étienne F. L. Raoul, surgeon and botanist on the French corvette L’Aube, author of Choix de plantes de la Nouvelle Zélande.
 Description

Sub-shrub or low shrub to 0.3 m tall. Stems ascending to erect, eglandular-pubescent; hairs uniform, short, reflexed, appressed; minute glandular hairs sometimes also present. Leaf bud indistinct, leaves separating while very small, opposite-decussate, erecto-patent to spreading or recurved, separating early; lamina coriaceous, usually oblanceolate to obovate or spathulate, sometimes linear-oblanceolate, 7–25 mm long, 2–9 mm wide, glossy green bronze-green or yellowish-green above, dull pale green to yellowish-green beneath; midrib evident; surfaces glabrous; margin glabrous or eglandular- and glandular-ciliate towards base, serrate or bluntly serrate, teeth in 2–6, rarely 0–1, sometimes up to 8, pairs; apex acute or sub-acute and usually sub-apiculate; base cuneate; petiole 2–5 mm long. Inflorescence a terminal compound spike or raceme, 20–55 mm long; flowers crowded, 16–100 per inflorescence, all bisexual; bracts opposite, becoming alternate, deltoid, > pedicels; pedicels erecto-patent, 0–0.5 mm long, eglandular-hairy all around. Calyx lobes 4, the anterior pair usually fused at least ⅔-way to apex, rarely less or almost free, obtuse, 2–4 mm long, unequal, eglandular-ciliate, or mixed eglandular- and glandular-ciliate. Corolla 6–8 mm diameter; tube pink, purplish or white, 2.0–3.5 mm long, about = calyx, glabrous; lobes 4, pink, purplish, or white, erect to spreading, sub-equal, lanceolate, elliptic, oblong or rhomboid, 2.5–3.5 mm long, sub-acute to obtuse; nectar guides absent. Stamen filaments white, 1.5–2.0 mm long; anthers yellow. Style glabrous, 4–6 mm long. Capsules turgid or weakly latiseptate, truncate, glabrous, 3–4 mm long, 1.8–3.0 mm at widest point. Seeds fusiform or irregular, flattened or weakly flattened, winged, smooth, straw-yellow to pale brown, 1.2–2.2 mm long.

 Recognition

Among the species of the sun hebe group, plants of V. hulkeana and V. lavaudiana can be distinguished from V. raoulii plants by their larger leaves.

V. maccaskillii plants are similar in usually having the anterior calyx lobes fused, but are distinguished by their almost divaricating and compact habit, shorter, more rounded leaves with fewer teeth, and usually a small fifth 5-calyx lobe.

V. scrupea plants are similar in having four calyx lobes, but their leaves are narrower, more toothed, and more acute, flowers are smaller with shorter stamens, and the anterior calyx lobes are free.

V. pentasepala plants may be similar in size and overall appearance, but are usually taller and more stiffly erect, and distinguished by having five calyx lobes that are all separate.

 Distribution

South Island: Marlborough (Seaward Kaikōura Range, Tarndale); Canterbury (Seaward Kaikōura Range, Mt Terako, Hanmer Springs region, Hurunui and Waiau Valleys, and along the foothills of the southern alps to Rakaia Gorge). I have accepted and mapped Norton and Molloy’s (2009) record from Mt Cass; those plants have broader leaves, and often the anterior calyx lobes there are free.

Two collections said to be from Banks Peninsula are regarded as doubtful and not accepted here.

 Habitat

Rock outcrops in scrub and grassland, occasionally on talus or scree. Recorded elevations range from 350 to 1282 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

Although the distribution of V. raoulii overlaps those of V. scrupea, V. hulkeana, V. maccaskillii, and V. pentasepala, wild hybrids have not been noted. Veronica ‘Hagley Park’ is the hybrid V. hulkeana × raoulii, which arose in cultivation (Metcalf 2001).

 Phenology

Flowers: September–December; fruits: December–February.

 Cytology

2n = 42 (Hair 1967, as Hebe raoulii var. raoulii).

 Notes

Veronica raoulii is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “sun hebe” group (Albach & Meudt 2010). Chloroplast DNA sequence data place V. raoulii close to V. hulkeana and V. lavaudiana; ITS sequence data indicate a relationship with V. pentasepala (Albach & Meudt 2010). A morphology-based cladogram (Garnock-Jones 1993) suggests a close relationship with V. maccaskillii.

Cultivars

Veronica ‘Hagley Park’ is the cultivated hybrid between V. hulkeana and V. raoulii.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Heliohebe raoulii (Hook.f.) Garn.-Jones] [Not Threatened]
Garnock-Jones, P.J. 1993: Heliohebe (Scrophulariaceae-Veroniceae), a new genus segregated from Hebe. New Zealand Journal of Botany 31: 323–339.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Garnock-Jones, P.J.; Lloyd, D.G. 2004: A taxonomic revision of Parahebe (Plantaginaceae) in New Zealand. New Zealand Journal of Botany 42: 181–232. [as Heliohebe raoulii (Hook.f.) Garn.-Jones; Parahebe raoulii (Hook.f.) Heads]
Hair, J.B. 1967: Contributions to a chromosome atlas of the New Zealand flora — 10. Hebe (Scrophulariaceae). New Zealand Journal of Botany 5: 322–352.
Heads, M.J. 1994: A biogeographic review of Parahebe (Scrophulariaceae). Botanical Journal of the Linnean Society 115: 65–89.
Hooker, J.D. 1864: Handbook of the New Zealand Flora: a systematic description of the native plants of New Zealand and the Chatham, Kermadec's, Lord Auckland's, Campbell's and Macquarie's Islands. Part I. Reeve, London.
Metcalf, L.J. 2001: International Register of Hebe cultivars. Royal New Zealand Institute of Horticulture, Lincoln, Canterbury.
Norton, D.A.; Molloy, B.P.J. 2009: Heliohebe maccaskillii (Plantaginaceae) — a new rank for a threatened limestone endemic, North Canterbury, New Zealand. New Zealand Journal of Botany 47: 405–409.