Classification
 Nomenclature
Scientific Name:
Veronica hulkeana F.Muell., Edinburgh New Philos. J. 14: 157 (1861)
Synonymy:
  • Hebe hulkeana (F.Muell.) Cockayne & Allan, Trans. New Zealand Inst. 57: 43 (1926)
  • Heliohebe hulkeana (F.Muell.) Garn.-Jones, New Zealand J. Bot. 31: 328 (1993)
  • Parahebe hulkeana (F.Muell.) Heads, Bot. J. Linn. Soc. 115: 82 (1994)
Lectotype (designated by Garnock-Jones 1993): MEL 1594290, Veronica hulkei, Melb. Bot. Gard. Jan 30th 1862
Etymology:
Mueller named the species after Mr T.H. Hulke of New Plymouth.
 Description

Shrub to 0.7 m tall. Stems decumbent to ascending, glabrous or puberulent; hairs uniform. Leaf bud indistinct; leaves separating while very small, opposite-decussate, erecto-patent to spreading; lamina coriaceous, lanceolate, ovate, or elliptic, 10–60 mm long, 5–35 mm wide, glossy green or dark green above, dull pale green to white or pinkish beneath; midrib evident, secondary veins weakly evident above; surfaces glabrous or with eglandular hairs along midrib above; margin glabrous or rarely ciliolate towards apex, bluntly to sharply serrate; teeth or lobes in 5–20 pairs; apex obtuse to acute; base cuneate to truncate; petiole 5–25 mm long. Inflorescence a terminal compound raceme, 90–480 mm long; flowers crowded, 50–1500, all bisexual; bracts opposite below, becoming alternate above, ovate to deltoid, > pedicels; pedicels spreading, 0–1 mm long, glabrous or glandular- to eglandular-puberulent all around. Calyx lobes 4, obtuse to acuminate, equal, 1.5–3.0 mm long, glandular- to eglandular-ciliate. Corolla 6–10 mm diameter; tube purplish, rarely white, 1.0–1.5 mm long, < calyx, glabrous; lobes 4, pale purplish, rarely white, unequal, spreading, ovate to obovate, 3–5 mm long, obtuse; nectar guides absent. Stamen filaments white or pale purplish, 1.0–1.5 mm long; anthers yellow. Style glabrous or eglandular-hairy, 3.0–4.5 mm long. Capsules turgid, obtuse to emarginate, usually glabrous, sometimes eglandular-hairy, rarely glandular-hairy, 2.5–4.5 mm long, 2–3 mm at widest point. Seeds fusiform to irregular, weakly flattened, winged, smooth, or wrinkled on back, pale to dark brown, 1.0–2.4 mm long.

 Key
1Young stems and petioles puberulent; smaller inflorescence branches densely puberulent or glandular-puberulent; calyx lobes usually obtuse, rarely sub-acute, closely eglandular-ciliatesubsp. hulkeana
Stems and petioles glabrous; inflorescence branches either glabrous or sparsely eglandular- or glandular-puberulent; calyx lobes acuminate or acute, distantly glandular-ciliolate or with mixed glandular and eglandular ciliasubsp. evestita
 Recognition

V. hulkeana belongs to the distinctive sun hebe group, which is characterised by toothed or crenate leaves, terminal paniculate inflorescences, style very much > stamens, ciliate nectarial disc, and fusiform seeds. Among that informal grouping, plants of V. hulkeana can be distinguished by their large leaves with glossy adaxial surfaces and their very large and many-flowered panicles. V. lavaudiana plants have quite large leaves, but these are dull, and their inflorescences have glandular hairs.

Unidentified plants collected from the Omaka River and Black Birch Creek, Marlborough (e.g., CHR 470178) resemble V. pentasepala in their erect habit and narrow leaves, but are more like V. hulkeana in their pale lilac flowers and four free calyx lobes, and like V. scrupea in their four free calyx lobes and narrow leaves that are sometimes acute.

 Distribution

South Island: Marlborough (in the east from near Taylor Pass southwards), Canterbury (in the east from Mt Grey northwards).

 Habitat

Rock outcrops, cliffs, gorges, and sometimes in alluvial gravel. Recorded elevations range from 5 to 1281 m.

 Biostatus
Indigenous (Endemic)
Number of subspecific taxa in New Zealand within Veronica hulkeana F.Muell.
CategoryNumber
Indigenous (Endemic)2
Total2
 Cytology

2n = 42 (Hair 1967, as Hebe hulkeana). The voucher for this record is identified as subsp. hulkeana.

 Notes

Veronica hulkeana is classified in V. subg. Pseudoveronica sect. Hebe and informally in the “sun hebe” group (Albach & Meudt 2010).

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe hulkeana (F.Muell.) Andersen]
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
Garnock-Jones, P.J. 1993: Heliohebe (Scrophulariaceae-Veroniceae), a new genus segregated from Hebe. New Zealand Journal of Botany 31: 323–339.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Garnock-Jones, P.J.; Lloyd, D.G. 2004: A taxonomic revision of Parahebe (Plantaginaceae) in New Zealand. New Zealand Journal of Botany 42: 181–232. [as Heliohebe hulkeana (F.Muell.) Garn.-Jones; Parahebe hulkeana (F.Muell.) Heads]
Hair, J.B. 1967: Contributions to a chromosome atlas of the New Zealand flora — 10. Hebe (Scrophulariaceae). New Zealand Journal of Botany 5: 322–352.
Heads, M.J. 1994: A biogeographic review of Parahebe (Scrophulariaceae). Botanical Journal of the Linnean Society 115: 65–89.
Mueller, F. 1861: Botanical Society of Edinburgh: Observation on some hitherto undescribed plants from New Zealand. Edinburgh New Philosophical Journal 14: 157–158.