Classification
 Nomenclature
Scientific Name:
Veronica notialis Garn.-Jones in Garnock-Jones et al., Taxon 56: 578 (2007)
Synonymy:
  • Hebe pauciflora G.Simpson & J.S.Thomson, Trans. & Proc. Roy. Soc. New Zealand 73: 166 (1943)
  • Leonohebe pauciflora (G.Simpson & J.S.Thomson) Heads, Bot. Soc. Otago Newsl. 5: 10 (1987)
Holotype: Kepler Range, near Fowler Pass, grassland in open situations, G. Simpson, Mar 1942, CHR 75689. Probable isotypes: CHR 97607, K, and AK 22903
Etymology:
Notialis: southern. This species is endemic to western Fiordland, making it one of the world’s southernmost veronicas.
 Description

Spreading low shrub to 0.2 m tall. Stems decumbent to erect, eglandular-pubescent or glabrous; hairs bifarious. Leaf bud distinct, its outer leaves fully grown, diverging but leaving a broad, acute sinus at the base. Leaves opposite-decussate, erecto-patent to spreading; lamina coriaceous, rigid, ovate to rhomboid or spathulate or orbicular, 2–6 mm long, 2–6 mm wide, glossy pale green or yellowish to dark green above and beneath, weakly keeled but veins not evident; surfaces glabrous; margins eglandular-pubescent with long, tangled and often branching hairs, sometimes glabrous or finely papillate, entire, translucent, cartilaginous; apex obtuse to sub-acute or bluntly sub-apiculate; base cuneate; petiole broad, 1–2 mm long. Inflorescence a lateral spike or raceme, 6–14 mm long; flowers crowded, 2–4 or sometimes solitary, all bisexual; bracts opposite-decussate, connate, lanceolate to deltoid, > pedicels; pedicels erecto-patent, 0–1 mm long, glabrous or eglandular-hairy all around. Calyx lobes 4, acute, 4–6 mm long, sub-equal, eglandular-ciliate with tangled branching hairs. Corolla 6–9 mm diameter; tube white, 3–4 mm long, ≥ calyx, glabrous; lobes 4, white, spreading to recurved, sub-equal, elliptic to orbicular, 3.0–4.5 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 1.5–2.0 mm long; anthers pink, magenta, or purplish. Style 1.8–3.0 mm long. Capsule angustiseptate, truncate to didymous, glabrous, 4.5–5.0 mm long, 4 mm at widest point. Seeds ellipsoid, weakly flattened, smooth, pale brown, 1.0–1.5 mm long.

 Recognition

Veronica notialis is a very distinctive species. In habit the plants resemble plants of the V. odora group and “Connatae”, but the angustiseptate capsules distinguish V. notialis. No other species is characterised by sinuous branching hairs on the leaf margins, although some whipcord hebes have very similar sinuate but unbranched hairs. The thick, translucent, cartilaginous cuticle on the leaves is also distinctive; through the dissecting microscope the leaf appears as if embedded in amber, especially at the margins.

Plants of V. murrellii and V. petriei have a similar habit to V. notialis plants, but differ in their softer dull or only slightly glossy leaves, which have a glabrous or minutely ciliolate margin, terminal, many-flowered inflorescences, narrower corolla lobes, and latiseptate acute to acuminate capsules.

 Distribution
 Habitat

Alpine grassland. Recorded elevations range from 1100 to 1500 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: December–March; fruits: January–March.

 Cytology

2n = 42 (see Bayly & Kellow 2006, as Hebe pauciflora).

 Notes

Veronica notialis is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Pauciflorae” (Albach & Meudt 2010; Bayly & Kellow 2006).

Chromosome number, broad, rounded corolla lobes, and angustiseptate capsules suggest V. notialis may be a lineage attached at a basal node within the New Zealand clade, and this is borne out by unpublished DNA sequence data collected by E.M. Low, which place it as sister to the rest of the shrubby hebe clade, close to the attachment of two other southern species with a similar overall appearance, V. petriei and V. murrellii .

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe pauciflora G.Simpson & J.S.Thomson] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe pauciflora G.Simpson & J.S.Thomson]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe pauciflora G.Simpson & J.S.Thomson] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Simpson, G.; Thomson, J.S. 1943: Notes on some New Zealand plants and descriptions of new species. Transactions and Proceedings of the Royal Society of New Zealand 73: 155–171.