Classification
 Nomenclature
Scientific Name:
Veronica dilatata (G.Simpson & J.S.Thomson) Garn.-Jones in Garnock-Jones et al., Taxon 56: 577 (2007)
Synonymy:
  • Hebe dilatata G.Simpson & J.S.Thomson, Trans. & Proc. Roy. Soc. New Zealand 73: 164 (1943)
Holotype: Blue Lake, Garvie Mountains, Otago, 1370 m above sea level, G. Simpson & J. S. Thomson, CHR 63426 (a collection mounted on three sheets, labelled 63426A, 63426B and 63426C)
  • = Hebe crawii Heads, Bot. Soc. Otago Newsl. 5: 11 (1987)
Holotype: Excelsior Peak, Takitimu Mountains, 4800', occasional spreading shrubs ca. 40 cm tall in fellfield, A. F. Mark, 2 Feb 1971, OTA 31283
Etymology:
The epithet dilatata might mean broadened or widened, from Latin dilatus, but Simpson and Thomson (1943) did not explain its derivation. Bayly and Kellow (2006) suggested it might refer to the spreading habit of some plants. However, Simpson and Thomson’s description mentions the corolla tube is widest at the throat, and it might thus be a reference to the dilated corolla tube.
 Description

Spreading low shrub to 0.4 m tall. Stems decumbent or ascending, eglandular-pubescent; hairs bifarious. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus acute. Leaves opposite-decussate to sub-distichous, erecto-patent to recurved; lamina coriaceous, obovate to spathulate to elliptic, 4–25 mm long, 4–11 mm wide, dull or somewhat glossy pale green or glaucescent above and beneath; midrib evident and two lateral veins sometimes faintly visible; surfaces glabrous or eglandular-hairy along midrib above; margin glabrous or minutely papillate, entire; apex obtuse to rounded, sometimes weakly plicate-emarginate; base abruptly cuneate; petiole 0.5–4.0 mm long. Inflorescence a lateral spike or raceme, tripartite or simple, 5–42 mm long; flowers crowded, 5–52, female or bisexual on separate plants, ⚥ > ♀; bracts opposite below and the lowermost usually flowerless, becoming alternate above, narrowly deltoid to lanceolate to elliptic to ovate, > pedicels; pedicels erecto-patent, 0.5–3.0 mm long, eglandular-hairy all around and sometimes also with short glandular hairs. Calyx lobes 4 or posterior small 5th lobe sometimes present, anterior pair fused ⅓ to completely, sub-acute to obtuse, 2.0–2.7 mm long, unequal, mixed glandular- and eglandular-ciliolate. Corolla 5–10 mm diameter; tube white, 1.2–2.5 mm long, ≤ calyx, glabrous; lobes 4, white, sub-erect to spreading, unequal, elliptic or oblong to ovate, 2.0–4.5 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 3.8–4.5 mm long (1.0–2.5 mm in staminodes); anthers magenta (pale in staminodes). Style glabrous, 3.4–7.0 mm long. Capsules latiseptate, sub-acute to obtuse, glabrous, 3.4–4.5 mm long, 2.1–2.8 mm wide. Seeds ovoid to subdiscoid, flattened, smooth, straw-yellow to pale brown, 0.9–1.1 mm long.

 Recognition

In Fiordland, west of the range of V. dilatata, two species, V. cockayneana and V. arganthera, have similar dull or glaucous leaves and a narrow, acute leaf bud sinus. In V. cockayneana the leaves are discolorous, glossy green above and dull glaucescent beneath; the plants tend to grow more erect but may be low-growing in exposed sites. V. arganthera plants are often larger (up to 1.5 m tall); that species is distinguished from V. dilatata by longer leaves (12–38 mm long, 5–11 mm wide), ciliate or ciliolate leaf margins, and white anthers. V. cockayneana​​​​​​​ plants have the same chromosome number as V. dilatata (2n = 120), whereas V. arganthera have 2n = 40.

 Distribution

South Island: South Otago and Southland (Eyre Mountains, Garvie Mountains, Umbrella Mountains, Takitimu Mountains). 

An early record from the Blue Mountains has not been supported by recent collecting.

 Habitat

Alpine tussock grassland, especially in rocky sites, rock outcrops, screes. Recorded elevations range from 823 to 1600 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: November–January, sometimes to March); fruits: November–March, persisting longer.

 Cytology

2n = 120 (see Bayly & Kellow 2006, as Hebe dilatata).

 Notes

Veronica dilatata is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Apertae” (small-leaved) (Albach & Meudt 2010; Bayly & Kellow 2006). Limited molecular studies have not provided strong evidence of close relationships of V. dilatata. Morphology and chromosome numbers suggest V. cockayneana might be a close relative, and beyond that it might be related to V. arganthera and to several similar species from Nelson (V. baylyi, V. rigidula, V. cryptomorpha, V. simulans).

Plants from the Takitimu Mountains tend to have pedicels > calyx, and they more often have some glandular hairs, than plants from the Garvie, Umbrella, and Eyre Mountains, which have pedicels ≤ calyx. They also have more tripartite inflorescences and their leaves are more glaucescent. These have been separated taxonomically (as Hebe crawii), but consistent and qualitative differences that would indicate the existence of genetically separate and differentiated lineages are lacking.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe crawii Heads; Hebe dilatata G.Simpson & J.S.Thomson]
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe dilatata G.Simpson & J.S.Thomson] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe dilatata G.Simpson & J.S.Thomson]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe dilatata G.Simpson & J.S.Thomson] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.
Simpson, G.; Thomson, J.S. 1943: Notes on some New Zealand plants and descriptions of new species. Transactions and Proceedings of the Royal Society of New Zealand 73: 155–171.