Classification
 Nomenclature
Scientific Name:
Veronica rigidula Cheeseman, Man. New Zealand Fl. 514 (1906)
Synonymy:
  • Hebe rigidula (Cheeseman) Cockayne & Allan, Trans. New Zealand Inst. 56: 20 (1926)
Lectotype (designated by Moore, in Allan 1961): Pelorus River, Marlborough, J. H. McMahon, AK 7919
Etymology:
In the protologue Cheeseman referred to the leaves as being rigid. Stearn (2004) translates rigidulus as “somewhat rigid”, and Bayly and Kellow (2006) relate this to the leaves.
 Description

Low, open-branched shrub to 0.6 m tall. Stems ascending to erect, eglandular-pubescent; hairs bifarious. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus acute. Leaves opposite-decussate, erecto-patent to spreading; lamina sub-coriaceous to coriaceous, lanceolate to narrowly or broadly elliptic to oblong to oblanceolate, 7–25 mm long, 2–9 mm wide, dull to slightly glossy green or yellowish-green above, dull glaucous to glaucescent beneath; midrib evident; surfaces glabrous or sometimes eglandular hairs along midrib above; margin glabrous, minutely papillate, entire; apex sub-acute to acute, acuminate to weakly plicate-acuminate; base cuneate; petiole 2.0–3.2 mm long. Inflorescence a lateral, usually tripartite, but sometimes simple or compound, raceme, 14–32 mm long; flowers crowded, 10–40, all bisexual; bracts opposite-decussate, sometimes becoming alternate above, ovate to deltoid, > pedicels; pedicels erecto-patent, 0.1–2.0 mm long, eglandular-hairy all around, or rarely absent. Calyx lobes 4, sub-acute to obtuse, 1.5–2.6 mm long, sub-equal, eglandular-ciliate or mixed glandular- and eglandular-ciliate. Corolla 5–7 mm diameter; tube white, 2.5–5.0 mm long, > calyx, glabrous; lobes 4, white, erecto-patent to spreading, becoming recurved, sub-equal, elliptic to ovate or rhomboid, 2–3 mm long, obtuse or sometimes posterior emarginate; nectar guides absent. Stamen filaments white, 3–4 mm long; anthers buff to pink. Style glabrous, 5–7 mm long. Capsules latiseptate, sub-acute to obtuse, glabrous, 2.4–4.0 mm long, 1.8–2.5 mm at widest point. Seeds ellipsoid to discoid or irregular, flattened, smooth, pale brown to brown, 0.8–1.6 mm long.

 Key
1Lamina concave when viewed from above, usually narrowly elliptic or sometimes elliptic or oblanceolate, mostly 3.6–4.4 times as long as broad; pedicels 0.1–0.8 mm longvar. rigidula
Lamina longitudinally folded and each half convex when viewed from above, elliptic to broadly elliptic, mostly 2.8–3.3 times as long as broad; pedicels 0.3–2.0 mm longvar. sulcata
 Recognition

Veronica rigidula is one of a few species characterised by discolorous leaves and usually tripartite inflorescences. V. rupicola plants are similar but differ in brighter green leaves, large bracts that almost completely obscure the larger calyx, sessile or sub-sessile flowers, and acuminate corolla lobes; its distribution is in eastern Marlborough. Plants of V. scopulorum, which grow near Raglan in the North Island, are also similar, and differ in the strongly tetragonous leaf buds with concave faces, and generally longer leaves and pedicels.

V. rigidula specimens have been confused in collections with V. subfulvid. V. subfulvida plants are similar because they also have a sinus, compound racemes and somewhat discolorous leaves, but they differ in their larger stature, leaves never glaucous (green to dark green above, paler beneath), and hairs inside the corolla tube.

 Distribution

South Island: Sounds Nelson.

 Habitat

Open rocky habitats, sparse scrub, outcrops, boulder falls, cliffs, and gorges. Recorded elevations range from 21 to 1524 m.

 Biostatus
Indigenous (Endemic)
Number of subspecific taxa in New Zealand within Veronica rigidula Cheeseman
CategoryNumber
Indigenous (Endemic)2
Total2
 Phenology

Flowers: November–February; fruits: December–March (persisting until October).

 Cytology

2n = 40 (see Bayly & Kellow 2006, as Hebe rigidula var. rigidula and var. sulcata).

 Notes

Veronica rigidula is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Apertae” (small-leaved) (Albach & Meudt 2010; Bayly & Kellow 2006). The phylogenetic position of Veronica rigidula is unclear. Similar species among the many that appear to be related to it on the basis of ITS sequence data (E.M. Low, unpublished) include V. rupicola and V. subfulvida.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Bayly, M.J.; Kellow, A.V.; Mitchell, K.A.; Markham, K.R.; de Lange, P.J.; Harper, G.E.; Garnock-Jones, P.J.; Brownsey, P.J. 2002: Descriptions and flavonoid chemistry of new taxa in Hebe sect. Subdistichae (Scrophulariaceae). New Zealand Journal of Botany 40(4): 571–602.
Cheeseman, T.F. 1906: Manual of the New Zealand Flora. Government Printer, Wellington.
Cockayne, L.; Allan, H.H. 1926: A proposed new botanical district for the New Zealand region. Transactions of the New Zealand Institute 56: 19–20.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.