The following generic description is modified from Zander (1993).
Plants yellow- or glaucous-green above, brown below, forming dense to lax interwoven mats on soil or rock. Stems irregularly branched, to 70 mm, in cross-section usually rounded-triangular, central strand absent or small, sclerodermis present. Leaves usually in 3 distinct ranks, erect-spreading to spreading when moist, usually appressed to incurved and flexuose when dry, triangular to ovate-triangular (in N.Z. species), acute to acuminate, margins variably recurved in basal ½ to ⅘ of leaf, entire to weakly serrulate (crenulate from bulging papillose cells in N.Z. species); upper laminal cells obscure, round-rhombic to quadrate, firm- to thick-walled, papillose; marginal cells not differentiated; lower laminal cells differentiated weakly in a very small group medially, or occasionally not differentiated. Costa strong, percurrent or occasionally failing before the apex; adaxial superficial cells elongate and smooth to weakly papillose; abaxial superficial cells papillose throughout; in cross-section with two stereid bands. Axillary hairs to 9 cells with basal 1–2 cells thick-walled. Laminal KOH colour reaction yellow (in N.Z. species) to orange.
Dioicous. Perichaetia terminal, or terminal and lateral in the same species and occasionally on the same plant, with inner leaves strongly sheathing the seta. Perigonia terminal and lateral in clusters (not seen in N.Z. material). Setae elongate. Capsules cylindric to ellipsoid. Operculum conic or rostrate. Peristome of 16 teeth, short, straight, variously cleft, lacking a basal cylinder. Spores 10–17 µm, papillose.
A genus of about 10 species, found in Australasia, South Africa, southern South America, western North America, and Europe; usually in Mediterranean climates. Molecular studies (Hedderson & Zander 2007) indicate a relationship with the genus Leptodontium, confirming the detailed morphological analyses of Zander (1993).
Two species occur in N.Z., both of which are Australasian in distribution.
1 | Shoots terete when dry and with leaves closely appressed; stems red; leaf margins decurrent on stem; laminal cells thick-walled, with 1–2 papillae per cell | T. papillata |
1' | Shoots catenulate when dry and with leaves flexuose-twisted; stems green to brown; leaf margins not decurrent on stem; laminal cells firm-walled with c. 4 papillae per cell | T. tasmanica |
Category | Number |
---|---|
Indigenous (Non-endemic) | 2 |
Total | 2 |