Classification
 Nomenclature
Scientific Name:
Veronica masoniae (L.B.Moore) Garn.-Jones in Garnock-Jones et al., Taxon 56: 578 (2007)
Synonymy:
  • Hebe pauciramosa var. masoniae L.B.Moore in Allan, Fl. New Zealand 1, 926 (1961) – as masonae
  • Leonohebe masoniae (L.B.Moore) Heads, Bot. Soc. Otago Newsl. 5: 10 (1987)
  • Hebe masoniae (L.B.Moore) Garn.-Jones, Austral. Syst. Bot. 6: 478 (1993)
Holotype: Head of Cobb Valley, Nelson, R. Mason, 23 Feb 1946, CHR 54435
  • = Leonohebe masoniae var. rotundata Heads, Bot. Soc. Otago Newsl. 5: 11 (1987)
Holotype: mineral belt north of Cobb Reservoir, tussockland, 3500 ft, A. P. Druce, Nov 1980, CHR 389212
Etymology:
Named after Ruth Mason (1913–1990), botanist, who collected the type material and noted some distinct features of the plants.
 Description

Shrub to 0.5 m tall. Stems decumbent or ascending to erect, eglandular-pubescent; hairs bifarious. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus broadly shield-shaped. Leaves opposite-decussate, erecto-patent to spreading; lamina coriaceous, rigid, oblong to broadly oblong, to elliptic to sub-orbicular, 3–10 mm long, 4–8 mm wide, glossy green to dark green above and beneath, sometimes yellowish, especially on margins; midrib beneath and occasionally two faint lateral veins evident; surfaces glabrous; margin smooth or minutely papillate distally, ciliolate when young, entire; apex obtuse and slightly apiculate; base truncate to sub-cordate; petiole 1–2 mm long. Inflorescence a simple or compound terminal spike, 8–18 mm long; flowers crowded, 2–14, all bisexual; bracts opposite-decussate and connate, elliptic to sub-orbicular or rhomboid, ≥ calyx; pedicels absent. Calyx lobes 4, obtuse to sub-acute, 5.0–6.5 mm long, unequal, eglandular-ciliate, rarely with a few glandular hairs as well. Corolla 10–15 mm diameter; tube white, 4.5–6.0 mm long, = or slightly > calyx, eglandular-hairy in tube, with small glandular hairs at throat; lobes 4, white, sometimes purplish or pink, erecto-patent to recurved, sub-equal, elliptic to broadly obovate or sub-orbicular, 5–7 mm long, rounded; nectar guides absent. Stamen filaments white 5–8 mm long; anthers magenta. Style glabrous, 7–9 mm long. Capsules latiseptate, sub-acute to obtuse, glabrous, 4–5 mm long, 3.5–4.0 mm at widest point. Seeds ellipsoid to circular, flattened, smooth, pale brown, 1.5–2.1 mm long.

 Recognition
Morphological features to distinguish Veronica odora and look-alikes. Species are ordered according to similarity of their habit.
 

odora

mooreae

pauciramosa

masoniae

venustula

brachysiphon

Leaf bud sinus

broad, shield-shaped

narrow & acute to broad & shield shaped

broad, shield-shaped

broad, shield-shaped

narrow, acute

narrow, acute

Leaf margin

sharply bevelled; glabrous

bevelled at 90º to surfaces, glabrous

rounded; glabrous or with minute hairs or denticles

rounded, papillate towards apex; ciliolate when young

weakly bevelled; ciliolate when young, becoming glabrous or papillate

weakly bevelled; ciliolate to ciliate when young, becoming papillate

Stomata

adaxial – (but often + at Arthur’s Pass); abaxial +

adaxial – (+ at Caswell Sound, Denniston); abaxial +

adaxial +; abaxial +

adaxial +; abaxial +

adaxial ±; abaxial +

adaxial ±; abaxial +

Midrib

sharply keeled beneath

depressed above; prominent beneath

rounded beneath and flattened just short of apex

keeled throughout

evident but not keeled

evident but not keeled

Inflorescence

terminal + usually lateral spikes

lateral spikes only

lateral spikes only

terminal spikes only

lateral raceme, sometimes ternate, rarely compound.

lateral raceme, sometimes ternate.

Bracts

not overtopping calyx

< calyx

< calyx

≥ calyx

≥ pedicels, < calyx

≥ pedicels, < calyx

Bracts and flowers

opposite

opposite

opposite

opposite

opposite below, becoming alternate

alternate, or lowermost opposite

Pedicels

0 mm

0–1 mm

0–0.5 mm

0 mm

0.5–7.0  mm

0.6–3.0 mm

Calyx, anterior lobes

free

free or fused to ⅓-way

fused > ⅔-way

free

free

free

Corolla lobes

narrow

broad

narrow

broad

± broad

± broad

 Distribution
 Habitat

Sub-alpine to penalpine tussock grassland, carpet grass (Chionochloa australis) and low scrub, sometimes in wet sites. Recorded elevations range from 769 to 1553 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: October–April; fruits: January–May (sometimes persisting until November).

 Cytology

2n = 118 (see Bayly & Kellow 2006, as Hebe masoniae).

 Notes

Veronica masoniae is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Buxifoliatae” (Albach & Meudt 2010; Bayly & Kellow 2006). Although V. masoniae resembles V. pauciramosa (with which it was included until 1987) and V. odora, analysis of ITS sequence data (E.M. Low, unpublished) places it among a large and poorly resolved clade of shrubby hebes that mostly have 2n = 40, 80, 120. Further research to clarify its position and origins is needed.

Flowers of both V. masoniae and V. pauciramosa have characteristic dark red stigmas.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington. [as Hebe masoniae (L.B.Moore) Garn.-Jones; Hebe pauciramosa var. masoniae L.B.Moore; Leonohebe masoniae (L.B.Moore) Heads var. masoniae]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe masoniae (L.B.Moore) Garn.-Jones] [Not Threatened]
Garnock-Jones, P.J. 1993: Phylogeny of the Hebe complex (Scrophulariaceae: Veroniceae). Australian Systematic Botany 6(5): 457–479.
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Heads, M. 1987: New names in New Zealand Scrophulariaceae. Botanical Society of Otago Newsletter 5: 4–11.