Classification
 Nomenclature
Scientific Name:
Veronica lanceolata Benth. in de Candolle, Prodr. 10 462 (1846)
Synonymy:
  • Veronica catarractae var. lanceolata (Benth.) Hook.f., Bot. Antarct. Voy. II. (Fl. Nov.-Zel.) Part I, 195 (1853)
  • Parahebe catarractae subsp. lanceolata (Benth.) Garn.-Jones in Garnock-Jones & Langer, New Zealand J. Bot. 18: 294 (1980)
  • Parahebe lanceolata (Benth.) Garn.-Jones in Garnock-Jones & Lloyd, New Zealand J. Bot. 42: 203 (2004)
Lectotype (designated by Garnock-Jones & Langer 1980): N. Zealand, Ms of the interior, Dieffenbach, K (sheet bearing two stems)
  • = Veronica diffusa Hook.f. in Hooker, Icon. Pl. 7, t. 645 (1844) nom. illeg., non Veronica diffusa Raf. 1838
  • Veronica catarractae var. diffusa Hook.f., Handb. New Zealand Fl. 216 (1864) nom. nov. pro Veronica diffusa Hook.f. 1844
  • Parahebe diffusa (Hook.f.) W.R.B.Oliv., Rec. Domin. Mus. 1: 230 (1944)
  • Parahebe catarractae subsp. diffusa (Hook.f.) Garn.-Jones, New Zealand J. Bot. 18: 292 (1980)
Holotype: K, On Tongariro, Bidwill
  • = Veronica lanceolata var. angustifolia Benth. in de Candolle, Prodr. 10 463 (1846)
Lectotype (designated by Garnock-Jones & Langer 1980): New Zealand, Ms of the interior, Dieffenbach, the sheet bearing one small-leaved specimen, K
  • = Veronica irrigans Kirk, Trans. New Zealand Inst. 2: 94 (1870)
Lectotype (designated by Garnock-Jones & Langer 1980): Thames Gold Fields, T. Kirk, WELT 41535
Etymology:
The epithet lanceolata is a reference to the leaf shape of the type specimen, which is a common leaf shape in this species.
 Description

Sub-shrub to 0.45 m tall. Stems prostrate to erect, eglandular-pubescent or glabrous; hairs bifarious, rarely uniform. Leaf bud indistinct; leaves separating while very small, opposite-decussate, or sub-distichous on prostrate stems, erecto-patent to spreading or recurved; lamina sub-coriaceous or coriaceous, linear, lanceolate, ovate, elliptic, orbicular, rhomboid, or deltoid, rarely obovate or oblanceolate, 5–100 mm long, 1.5–25.0 mm wide, dull or glossy pale to dark green above, dull pale green or green or pinkish beneath; midrib and sometimes secondary veins evident; surfaces glabrous, sometimes with eglandular hairs along midrib above; margins glabrous, serrate; teeth in 1–15 pairs; apex sub-acute to acuminate, rarely obtuse; base cuneate, truncate, or sub-cordate; petiole 1–6 mm long. Inflorescence a lateral raceme, 30–350 mm long; flowers distant, 4–30, all bisexual; bracts alternate, linear, lanceolate, elliptic, ovate, or narrowly deltoid, < pedicels; pedicels erecto-patent, incurved at fruiting, 5–25 mm long, eglandular-hairy all around, rarely glabrous or glandular-hairy. Calyx lobes 4, obtuse to acuminate, 2–4 mm long, sub-equal, glandular- and/or eglandular-ciliate, occasionally pubescent on abaxial surfaces. Corolla 8–15 mm diameter; tube white and yellow, 0.5–1.5 mm long, < calyx, eglandular-hairy inside; lobes white, usually 4 or rarely 5 by division of posterior lobe, erecto-patent to spreading, unequal, narrowly oblong or elliptic to orbicular, 3.5–8.0 mm long, rounded, sometimes posterior emarginate; nectar guides magenta. Stamen filaments white, 2–4 mm long; anthers pink or magenta. Style glabrous, 3.5–7.0 mm long. Capsules angustiseptate to turgid, emarginate, glabrous, 2.5–4.5 mm long, 2.5–4.0 mm at widest point. Seeds discoid to obovoid, flattened, smooth, straw-yellow to brown, 0.5–1.1 mm long.

 Recognition

V. lanceolata is a highly variable species, and some of the regional variants can be hard to distinguish from other species in the speedwell hebe group. 

Large plants with coarsely toothed leaves may resemble V. catarractae, but that species is endemic to Fiordland and can be distinguished by its white leaf undersides and pedicel hairs in a single row. 

Smaller low-growing plants with broader leaves can resemble V. melanocaulon, which is endemic to Marlborough and can be distinguished by usually obovate or oblanceolate leaves, usually glabrous or sometimes glandular inflorescences, and very dark purplish-black stems that contrast with the pale green petioles. 

Small plants with short, broad leaves may resemble V. lyallii, but V. lyallii plants have shorter inflorescences, more rounded leaf teeth, and markedly arcuate, antrorse stem hairs; sometimes their flowers lack nectar guides and their leaves can be bronze-green. V. lyallii is confined to the South Island.

 Distribution

North Island: Auckland (Coromandel Peninsula, Pirongia), Taranaki from Herangi Range to Kaitake Range), Gisborne, Volcanic Plateau, Southern North Island.

South Island: Western Nelson (Wakamarama Range only).

 Habitat

Rock outcrops, especially on stream and river banks, rooted in silt in crevices, also on cliffs, ledges, and screes. Recorded elevations range from 10 to 1708 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

Garnock-Jones & Lloyd (2004) considered that plants with broad leaves and densely glandular inflorescences from mountains of the Volcanic Plateau might be the result of introgression with V. hookeriana.  Similarly, plants on Mt Hikurangi and the Maungaharuru Range appear to intergrade with local forms of V. hookeriana to some extent.

 Phenology

Flowers: November–March; fruits: January–May, and old fruits present year round.

 Cytology

2n = 42 (Garnock-Jones & Langer 1980, as Parahebe catarractae). Hair reported the same number (as Parahebe catarractae) from a cultivated plant of unknown origin and a plant from Mt Ruapehu that I have identified as V. hookeriana × lanceolata.

 Notes

Veronica lanceolata is classified in V. subg. Pseudoveronica sect. Hebe (Albach & Meudt 2010). Molecular phylogenetics (Albach & Meudt 2010) placed V. lanceolata firmly within the speedwell hebe clade, along with other species of Veronica that are characterised by lax inflorescences, short corolla tubes, nectar guides, plicate lateral corolla lobes, attenuate stamen filament bases, and turgid capsules, but the internal relationships within that clade are not clearly established.

Garnock-Jones & Lloyd (2004) described nine regional races in detail. These were not treated formally as subspecies or varieties because they are not clear-cut, tend to vary within and among populations, and overlap with each other in their morphological features.

(See: Regional variation in Veronica lanceolata​​​​​​​)

Regional variation in Veronica lanceolata.

Region

Stems

Leaf shape & size (mm)

Leaf apex

Leaf base

Pairs of teeth

Flower number

Inflorescence hairs

Bracts

Pedicel length & hairs

Calyx hairs

Coromandel, Taranaki, Paturau (NW Nelson)

ascending to erect; internodes 5–25 mm long; bifarious (rarely glabrous)

linear to lanceolate, 20–100 x 1.5–12 mm

acute to acuminate

cuneate

(3–) 6–15

9–30

glabrous or eglandular

linear or lanceolate, glabrous or ciliate

5–13 mm, eglandular or glandular all around

glabrous or ciliate

N Taranaki, Whanganui mudstone

stout, decumbent to erect; internodes 10–90 mm long; glabrous (rarely bifarious)

lanceolate, ovate, or elliptic, 15–80 x 8–35 mm

acute to acuminate

cuneate to cordate

(7–) 10–18

20–35

glabrous or eglandular

lanceolate or narrow elliptic, glabrous or ciliate

8–18 mm, eglandular all around or sometimes glandular

margins or adaxial surface, eglandular or glandular

Volcanic Plateau

slender to stout, decumbent to erect; internodes 5–70 mm long; bifarious or glabrous

lanceolate, ovate, or elliptic, 8–45 x 3–25 mm

acute or subacute

cuneate to cordate

(3–) 6–10 

8–30

eglandular sometimes mixed with glandular

lanceolate, elliptic, or ovate

8–20 mm, eglandular and/or glandular all around

glabrous, or marginal, or abaxial, eglandular or glandular

East Cape, Kaimanawa, Kaweka, and Ruahine Ranges

slender to stout, prostrate to erect; internodes 5–70 mm long; bifarious, or rarely uniform

lanceolate to oblanceolate, elliptic, or ovate to obovate, 3–50 x 1.5–22 mm

subacute to acuminate

cuneate or subcordate

(2–) 4–9 (–12)

5–25

eglandular, rarely some glandular hairs (Maungaharuru Range)

lanceolate to narrowly elliptic

8–25 mm, eglandular all around (rarely some glandular (Maungaharuru Range))

glabrous or eglandular on margins

Lowland Hawke’s Bay & eastern Bay of Plenty

stout, ascending to erect; internodes 5–40 mm long, bifarious

lanceolate or elliptic, 5–55 x 3–15 mm

subacute to acute

cuneate

4–11

5–25

eglandular, sometimes also glandular; peduncle may be glabrous

lanceolate or elliptic

7–16 mm, eglandular or mixed eglandular & glandular all around

glabrous or eglandular on margins

Tararua & Aorangi Ranges (alpine)

moderately stout, decumbent to ascending; internodes 5–25 mm long, bifarious

ovate or elliptic, rarely lanceolate, 10–25 x 5–20 mm

subacute

subcordate to cordate, rarely abruptly cuneate

(3–) 6–12

8–20

eglandular and glandular

lanceolate or elliptic

6–9 (–11) mm, eglandular or glandular all around

glabrous or eglandular on margins

Tararua Range (lowland); similar plants at Kaituna R., NW Nelson

slender to stout, ascending to erect; internodes 3–35 mm long, bifarious

linear, lanceolate, oblanceolate, or elliptic, 7–40 x 2–13 mm

narrowly acute to subacute

cuneate

3–8 (–12)

6–20

eglandular, sometimes also glandular; peduncle may be glabrous

lanceolate, rarely elliptic

(5–) 8–12 mm, eglandular or mixed eglandular & glandular all around

marginal, eglandular or mixed eglandular & glandular, rarely glabrous

Southern Hawke’s Bay & lowland Wairarapa

very slender prostrate to decumbent; internodes 5–30 mm long, bifarious

lanceolate or oblanceolate to broadly elliptic, 5–20 x 2.5–10 mm

subacute or acute

cuneate

2–6

4–8 (–12)

eglandular (peduncle glabrous)

elliptic

6–10 mm, eglandular all around, rarely glabrous

marginal eglandular, or glabrous

Rimutaka Range & Cape Palliser

stout, ascending to erect; internodes 8–70 mm long, bifarious

broadly ovate or elliptic, 8–30 x 7–25 mm

obtuse to subacute

cuneate, truncate, or cordate

(4–) 6–9 (–12)

6–15 (–30)

mixed eglandular & glandular, peduncle glabrous

elliptic

8–12 mm, eglandular all around (glandular at C. Palliser)

glabrous or rarely 1–2 hairs at apex

 Bibliography
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de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened]
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