Classification
 Nomenclature
Scientific Name:
Veronica chathamica Buchanan, Trans. New Zealand Inst. 7: 338 (1875)
Synonymy:
  • Hebe chathamica (Buchanan) Cockayne & Allan, Trans. New Zealand Inst. 57: 22 (1926)
Lectotype (designated by Bayly & Kellow 2006): Transactions of the New Zealand Institute 7: 338, plate 13, fig. 1 (1875)
  • = Veronica coxiana Kirk, Trans. New Zealand Inst. 28: 529 (1896)
  • Veronica chathamica var. coxiana (Kirk) Cheeseman, Man. New Zealand Fl., ed. 2, 794 (1925)
  • Hebe coxiana (Kirk) Cockayne, Trans. New Zealand Inst. 60: 470 (1929)
Lectotype (designated by Moore, in Allan 1961): Chatham Islands, F. A. D. Cox, WELT 5295
Etymology:
The epithet chathamica refers to its distribution on the Chatham Is.
 Description

Spreading low or mat-forming shrub to 0.25 m tall. Stems prostrate to decumbent, or pendent on cliffs, eglandular-pubescent; hairs uniform or rarely bifarious. Leaf bud indistinct; leaves separating while small, opposite-decussate to sub-distichous, erecto-patent to recurved; sinus absent or very rarely minute and rounded; lamina coriaceous, elliptic to obovate or oblanceolate, 8–33 mm long, 3–17 mm wide, dull green to dark green above, green beneath; midrib evident; surfaces with eglandular hairs on midrib, or eglandular-hairy, or rarely glabrous; margin pubescent or glabrous, entire; apex obtuse to sub-acute; base cuneate; petiole absent. Inflorescence a lateral raceme, 13–41 mm long; flowers crowded, 20–40, all bisexual; bracts alternate or the lowest pair opposite, lanceolate or linear-lanceolate, ≥ pedicels; pedicels erecto-patent, 1–2.6 mm long, eglandular-hairy all around. Calyx lobes 4, acute to acuminate, rarely sub-acute, 1.7–3.5 mm long, sub-equal, eglandular-ciliate or with a few glandular hairs as well, often also hairy on outer surface. Corolla 6–8 mm diameter; tube white, 2.5–4.0 mm long, > calyx, eglandular-hairy inside and at bases of lobes; lobes 4, white or tinged pale purple, erecto-patent to spreading, sub-equal, elliptic to ovate, 2.0–3.5 mm long, obtuse to rounded; nectar guides absent. Stamen filaments white, 4.0–4.5 mm long; anthers pale brown to pale purple. Style glabrous or eglandular-hairy, 5–6 mm long. Capsules latiseptate, sub-acute to obtuse, glabrous or eglandular-hairy, 3.5–5.0 mm long, 2.5–3.5 mm at widest point. Seeds ellipsoid to sub-discoid, flattened, smooth, brown, 1.2–1.6 mm long.

 Recognition

Veronica chathamica plants have smaller leaves than plants of the other two woody hebes on the Chatham Is., and their leaves are relatively broader in shape. V. dieffenbachii plants might be confused with them, but tend to have larger leaves, longer inflorescences with more flowers, and often a taller growth form.

 Distribution

Chatham Is. (The Sisters [Rangitatahi], Chatham I., Pitt I. [Rangiauria], South East I. [Rangatira]).

 Habitat

Coastal rocks and cliffs, low vegetation at beach and lagoon margins. Recorded elevations range from 0 to 100 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

Where V. chathamica and V. dieffenbachii occur together they may be hard to distinguish and it is possible that hybridisation is occurring.

 Phenology

Flowers: December–March; fruits: January–March, persisting all year.

 Cytology

2n = 40 (see Bayly & Kellow 2006, as Hebe chathamica).

 Notes

Veronica chathamica is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Occlusae” (Albach & Meudt 2010; Bayly & Kellow 2006).

V. chathamica is variable in a number of features: leaf shape and size, stem and leaf indumentum, and the shape and hairiness of the calyx. Some plants of V. dieffenbachii and V. chathamica have been observed with blue or purple pollen, but more often it is whitish.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Buchanan, J. 1875: On the Flowering Plants and Ferns of the Chatham Islands. Transactions of the New Zealand Institute 7: 333–341.
Cheeseman, T.F. 1925: Manual of the New Zealand Flora. Edition 2. Government Printer, Wellington.
Cockayne, L. 1929: New combinations in the genus Hebe. Transactions of the New Zealand Institute 60: 465–472.
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe chathamica (Buchanan) Cockayne & Allan] [Naturally uncommon]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe chathamica (Buchanan) Cockayne & Allan]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Naturally Uncommon]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe chathamica (Buchanan) Cockayne & Allan] [Naturally Uncommon]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Kirk, T. 1896: Notes on certain Veronicas, and descriptions of new species. Transactions of the New Zealand Institute 28: 515–531.