Classification
 Nomenclature
Scientific Name:
Veronica brachysiphon (Summerh.) Bean, Bull. Misc. Inform. Kew 1934: 224 (1934)
Synonymy:
  • Veronica traversii Hook.f., Curtis’s Botanical Magazine 104, Plate 6390 (1878) nom. illeg., non Veronica traversii Hook.f. 1864
  • Hebe brachysiphon Summerh., Kew Bull. 1927: 397 (1927)
Lectotype (designated by Bayly & Kellow 2004b): from Sir J. D. Hooker’s garden, 26 June 1893, K, flowering pieces on top left and bottom right only (these are mounted on the same sheet as pieces collected in March 1893, and another specimen – comprising two pieces – collected at Edinburgh Botanical Gardens)
Etymology:
The epithet brachysiphon means short tube, a reference to the corolla tube in relation to that of V. traversii, with which it had been confused.
 Description

Bushy and often rounded shrub, to 1.8 m tall. Stems erect; eglandular-pubescent with minute glandular hairs; hairs usually bifarious, sometimes uniform. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus narrow, acute. Leaves opposite-decussate to weakly sub-distichous, erecto-patent to spreading; lamina sub-coriaceous to rigid, ovate to obovate, usually elliptic, or sometimes oblanceolate or oblong, 5.5–25.5 mm long, 3.3–10.0 mm wide, usually more or less glossy or sometimes dull light to dark green above, paler and duller beneath; midrib and sometimes two lateral veins evident; surfaces glabrous except for eglandular hairs along midrib above; margin eglandular- and/or glandular-ciliolate when young, becoming minutely papillate with age, entire; apex sub-acute to obtuse, mostly acute, keeled beneath and weakly apiculate; base cuneate; petiole 0.5–3.5 mm long. Inflorescence a lateral raceme or sometimes ternate, 12–41 mm long; flowers crowded, 9–36, female or bisexual on separate plants, ⚥ > ♀; bracts alternate or the lowest opposite, lanceolate to ovate or narrowly deltoid, ≥ pedicels; pedicels erecto-patent, 0.6–3.0 mm long, eglandular-hairy all around or sometimes almost glabrous. Calyx lobes 4–5 (5th small, posterior), obtuse to sub-acute, occasionally emarginate, 1.9–3.0 mm long, equal or sub-equal, glandular- or mixed glandular- and eglandular-ciliolate. Corolla 6–8 mm diameter; tube white, 1.6–4.0 mm long, > calyx, eglandular-hairy inside; lobes 4, white or sometimes pale purplish, erecto-patent to spreading, sub-equal, usually elliptic to ovate, sometimes oblong, obovate, or orbicular; nectar guides absent. Stamen filaments white, 1.0–4.5 mm long; anthers magenta to purplish. Style glabrous, 4.2–7.2 mm long. Capsules latiseptate, sub-acute, usually glabrous or rarely eglandular hairy, 3–6 mm long, 2.3–4.5 mm at widest point. Seeds ellipsoid to oblong, flattened, weakly wrinkled and undulate at margin, pale brown to brown, 0.8–2.8 mm long.

 Recognition
Morphological features to distinguish Veronica odora and look-alikes. Species are ordered according to similarity of their habit.
 

odora

mooreae

pauciramosa

masoniae

venustula

brachysiphon

Leaf bud sinus

broad, shield-shaped

narrow & acute to broad & shield shaped

broad, shield-shaped

broad, shield-shaped

narrow, acute

narrow, acute

Leaf margin

sharply bevelled; glabrous

bevelled at 90º to surfaces, glabrous

rounded; glabrous or with minute hairs or denticles

rounded, papillate towards apex; ciliolate when young

weakly bevelled; ciliolate when young, becoming glabrous or papillate

weakly bevelled; ciliolate to ciliate when young, becoming papillate

Stomata

adaxial – (but often + at Arthur’s Pass); abaxial +

adaxial – (+ at Caswell Sound, Denniston); abaxial +

adaxial +; abaxial +

adaxial +; abaxial +

adaxial ±; abaxial +

adaxial ±; abaxial +

Midrib

sharply keeled beneath

depressed above; prominent beneath

rounded beneath and flattened just short of apex

keeled throughout

evident but not keeled

evident but not keeled

Inflorescence

terminal + usually lateral spikes

lateral spikes only

lateral spikes only

terminal spikes only

lateral raceme, sometimes ternate, rarely compound.

lateral raceme, sometimes ternate.

Bracts

not overtopping calyx

< calyx

< calyx

≥ calyx

≥ pedicels, < calyx

≥ pedicels, < calyx

Bracts and flowers

opposite

opposite

opposite

opposite

opposite below, becoming alternate

alternate, or lowermost opposite

Pedicels

0 mm

0–1 mm

0–0.5 mm

0 mm

0.5–7.0  mm

0.6–3.0 mm

Calyx, anterior lobes

free

free or fused to ⅓-way

fused > ⅔-way

free

free

free

Corolla lobes

narrow

broad

narrow

broad

± broad

± broad

 Distribution

South Island: Sounds Nelson (Red Hills Range), Marlborough, Canterbury, Westland, from the Main Divide to the eastern foothills, reaching a southern limit near Mt Hutt.

 Habitat

Montane to sub-alpine tussock grassland and scrub and in beech forest close to tree line. Recorded elevations range from 600 to 1605 m.

 Biostatus
Indigenous (Endemic)
 Hybridisation

The difficulty in distinguishing V. brachysiphon from V. subfulvida might be due to introgression.

 Phenology

Flowers: October–March; fruits: January–May, persisting until December.

 Cytology

2n = 120 (see Bayly & Kellow 2006, as Hebe brachysiphon).

 Notes

Veronica brachysiphon is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Apertae” (small-leaved) (Albach & Meudt 2010; Bayly & Kellow 2006).

The seed description above is based on very few specimens and might understate the variability.

 Bibliography
Bayly, M.J.; Kellow, A.V. 2004: Lectotypification of names of New Zealand members of Veronica and Hebe (Plantaginaceae). Tuhinga, Records of the Museum of New Zealand Te Papa Tongarewa 15: 43–52.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Bean, W.J. 1934: Miscellaneous Notes. Bulletin of Miscellaneous Information, Royal Gardens, Kew 1934(5): 219–232.
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Not Threatened]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe brachysiphon Summerh.] [Not Threatened]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Hooker, W.J. 1878: Curtis’s Botanical Magazine. Vol. 104. S.Curtis, London.
Summerhayes, V.S. 1927: The Type of Veronica Traversii. Kew Bulletin 1927: 395–398.