Classification
 Nomenclature
Scientific Name:
Veronica adamsii Cheeseman, Man. New Zealand Fl., ed. 2, 786 (1925)
Synonymy:
  • Hebe adamsii (Cheeseman) Cockayne & Allan, Trans. New Zealand Inst. 57: 15 (1926)
Lectotype (designated by Garnock-Jones & Clarkson 1994): Kapowairua to Tom Bowline’s Bay, North Cape, T. F. C[heeseman], Jan 1896, 1545 to Kew, AK 7666. Isolectotypes: AK 7665, 20330, WELT 79581 (although the wording of locality details on the label of this last sheet differs slightly from that on the lectotype)
Etymology:
Named after James Adams (1839—1906), former headmaster of Thames High School, who accompanied T.F. Cheeseman on his North Cape expedition in 1896.
Vernacular Name(s):
kōkōmuka; koromiko
 Description

Low spreading or bushy shrub to 1 m tall. Stems ascending to erect, glabrous to minutely eglandular-puberulent; hairs bifarious to uniform. Leaf bud distinct, its leaves appressed at margins until fully grown; sinus elliptic, acute. Leaves opposite-decussate, erecto-patent to spreading; lamina coriaceous, lanceolate to broadly lanceolate or narrowly elliptic to elliptic, 15–85 mm long, 5–18 mm wide, glossy above, dull beneath, bronze-green to yellowish-green; midrib evident, secondary veins faint; surfaces glabrous or sometimes with minute eglandular hairs along midrib above, or rarely sparse sessile glandular hairs beneath; margin glabrous or sparsely ciliolate, entire; apex sub-acute to acute; base cuneate; petiole 1–2 mm long. Inflorescence a lateral raceme, 45–150 mm long; flowers crowded, 20–65, all bisexual; bracts alternate, linear, < or rarely = pedicels; pedicels erecto-patent to spreading, 2.5–6.0 mm long, minutely puberulent all around. Calyx lobes 4, acuminate or sometimes the posterior pair rounded, 2.5–4.0 mm long, sub-equal, glabrous or with sparse, minute sessile glands on faces; mixed glandular- and eglandular-ciliolate. Corolla 7–8 mm diameter; tube white, 2.3–4.0 mm long, slightly < to slightly > calyx, eglandular hairy inside; lobes 4, white to purplish, fading white, erect to erecto-patent, sub-equal, ovate or elliptic to deltoid, 3.5–5.5 mm long, sub-acute; nectar guides absent. Stamen filaments white to purplish, 6.0–9.5 mm long; anthers mauve to purple. Style glabrous, 4.0–8.5 mm long. Capsules latiseptate, sub-acute to obtuse, glabrous, 3.4–6.0 mm long, 2.8–4.0 mm at widest point. Seeds ellipsoid to discoid, flattened, smooth, pale brown to brown, 1.4–2.5 mm long.

 Recognition

The presence of a sinus in the leafy bud is distinctive among large-leaved hebes of the mainland of Northland, shared only with V. diosmifolia (characterised by smaller incised leaves and compound inflorescences) and V. speciosa (characterised by broad, fleshy leaves and robust red flowers). 

Plants of V. adamsii could be mistaken for the similar V. perbella and V. saxicola; these all have large flowers with broad, acute or sub-acute corolla lobes. V. perbella and V. saxicola plants differ by having no sinus in the leaf bud, glabrous stems (rarely bifarious-puberulent in V. saxicola), and 2n = 40 chromosomes. Also, V. perbella capsules usually have a few short hairs along the septal groove and V. saxicola leaves are not glossy; both occur much further south than the known distribution range of V. adamsii.

Comparison of Veronica adamsii, V. macrocarpa, V. perbella, and V. saxicola, 4 similar species that all occur in Northland. This table is based on a more detailed one presented by de Lange P.J. & J.R. Rolfe (2008), New Zealand Journal of Botany 46: 531–545 (as Hebe).
 

adamsii

macrocarpa

perbella

saxicola

Stem hairs

uniformly to bifariously puberulent or glabrous

uniformly to bifariously puberulent

glabrous

usually glabrous or rarely bifarious-puberulent

Leaf bud sinus

present

absent

absent

absent

Lamina (above)

glossy

glossy

glossy

dull

Calyx colour

dark green or bronze-green

yellow-green

pink, reddish, violet, or red- to brown-green

yellowish- to dark green

Calyx lobes outer surface

minute sessile glands

glabrous or eglandular-puberulent

minute sessile glands

minute sessile glands

Corolla colour

white to mauve, rarely violet

white, rarely lilac, sometimes tinged lilac or pink

violet-red, violet, pink, deep mauve, very rarely carmine

pale lavender (rarely lilac)

Corolla tube

≤ calyx; hairy inside

> calyx; hairy inside

≤ calyx; hairy inside

< calyx; glabrous inside

Corolla lobes

subacute

obtuse to rounded

acute

subacute or bluntly acute

Capsule

glabrous

glabrous

glabrous except for a few short hairs along septal grooves

glabrous

Distribution

far North, known only from four sites, all within 1 km2, west of Spirits Bay

southern Northland from Whangārei and southern Kaipara Harbour southwards to South Auckland, Coromandel, and near Kawhia

western Northland between Ahipara and Waima Forest

known only from Maungaraho Rock, near Dargaville

 Distribution

North Island: Northland. Known only from Unuwhao Ridge, The Pinnacle, Pinnacle Ridge, and Tarure Hill, east of Spirits Bay.

 Habitat

East-facing breccia cliffs, usually among Astelia banksii or in vertical joints in the rock, sometimes in scrub. Recorded elevations range from 200 to 279 m.

 Biostatus
Indigenous (Endemic)
 Phenology

Flowers: October–July; fruits: April–August and probably earlier.

 Cytology

2n = 80 (see Bayly & Kellow 2006, as Hebe adamsii).

 Notes

Veronica adamsii is classified in V. subg. Pseudoveronica sect. Hebe and the informal group “Apertae” (large-leaved) (Albach & Meudt 2010; Bayly & Kellow 2006).

Plants from Tarure Hill have longer and wider leaves and longer, more robust inflorescences than plants in other populations (de Lange 1991, Garnock-Jones & Clarkson 1994). The Māori names koromiko, kōkōmuka, and variants – applied to V. stricta and V. salicifolia – might be applied also to similar large-leaved hebes, such as V. adamsii.

 Bibliography
Albach, D.C.; Meudt, H.M. 2010: Phylogeny of Veronica in the Southern and Northern Hemispheres based on plastid, nuclear ribosomal and nuclear low-copy DNA. Molecular Phylogenetics and Evolution 54: 457–471.
Bayly, M.J.; Kellow, A.V. 2006: An Illustrated Guide to New Zealand Hebes. Te Papa Press, Wellington.
Cheeseman, T.F. 1925: Manual of the New Zealand Flora. Edition 2. Government Printer, Wellington.
Cockayne, L.; Allan, H.H. 1926: The present taxonomic status of the New Zealand species of Hebe. Transactions of the New Zealand Institute 57: 11–47.
de Lange, P.J. 1991: Hebe `Unuwhao': notes on its distribution, ecology and conservation status. New Zealand Botanical Society Newsletter 24: 7–8.
de Lange, P.J.; Norton, D.A.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Hitchmough, R.; Townsend, A.J. 2009: Threatened and uncommon plants of New Zealand (2008 revision). New Zealand Journal of Botany 47: 61–96. [as Hebe adamsii (Cheeseman) Cockayne & Allan] [Nationally Critical]
de Lange, P.J.; Norton, D.A.; Heenan, P.B.; Courtney, S.P.; Molloy, B.P.J.; Ogle, C.C.; Rance, B.D. 2004: Threatened and uncommon plants of New Zealand. New Zealand Journal of Botany 42(1): 45–76. [as Hebe adamsii (Cheeseman) Cockayne & Allan]
de Lange, P.J.; Rolfe, J.R.; Barkla J.W.; Courtney, S.P.; Champion, P.D.; Perrie, L.R.; Beadel, S.N.; Ford, K.A.; Breitwieser, I.; Schönberger, I.; Hindmarsh-Walls, R.; Heenan, P.B.; Ladley, K. 2018: Conservation status of New Zealand indigenous vascular plants, 2017. New Zealand Threat Classification Series. No. 22. [Nationally Critical]
de Lange, P.J.; Rolfe, J.R.; Champion, P.D.; Courtney, S.P.; Heenan, P.B.; Barkla, J.W.; Cameron, E.K.; Norton, D.A.; Hitchmough, R.A. 2013: Conservation status of New Zealand indigenous vascular plants, 2012. New Zealand Threat Classification Series 3. Department of Conservation, Wellington. [as Hebe adamsii (Cheeseman) Cockayne & Allan] [Nationally Critical]
Garnock-Jones, P.J. 2023: Veronica. In: Breitwieser, I. (ed.) Flora of New Zealand – Seed Plants. Fascicle 9. Manaaki Whenua Press, Lincoln.
Garnock-Jones, P.J.; Albach, D.; Briggs, B.G. 2007: Botanical names in Southern Hemisphere Veronica (Plantaginaceae): sect. Detzneria, sect. Hebe, and sect. Labiatoides. Taxon 56: 571–582.
Garnock-Jones, P.J.; Clarkson, B. D. 1994: Hebe adamsii and H. murrellii (Scrophulariaceae) reinstated. New Zealand Journal of Botany 32(1): 11–15.