Plants small to medium-sized, occasionally robust. Stems erect, mostly lacking a central strand (but present in Dicranoweisia), with firm-walled cortical cells. Leaves mostly lanceolate, often strongly contorted when dry. Upper laminal cells short, often with some oblate, smooth or papillose, sometimes bearing longitudinal cuticular striations that extend over the cell walls (present in all N.Z. taxa).
Autoicous. Perigonia often on short branches at the base of the perichaetia. Setae elongate or short; capsules mostly erect, symmetric, and short-cylindric to ellipsoid, less often asymmetric, sometimes furrowed when dry; operculum rostrate. Peristome single or sometimes absent, with 16 teeth mostly undivided, variously ornamented. Calyptra cucullate, smooth.
There is little consensus concerning the size or the limits of this morphologically diverse family. The brief description above is derived primarily from the type genus and the genera occurring in N.Z. The type genus, the predominantly northern hemisphere Rhabdoweisia, was placed (with Amphidium) in a subfamily Rhabdoweisioideae within the Dicranaceae by Brotherus (1924). Modern Floras have either recognised that subfamily at the family rank (Smith 2004) or subsumed it within the larger Dicranaceae (e.g., Crum & Anderson 1981; Ireland et al. 1994). While retaining it in the Dicranales, Goffinet et al. (2009) presented a much expanded interpretation of the family, which is largely adopted here. Only three of the 15 genera they include in the family occur in N.Z. (They also included Kiaeria in the Rhabdoweisiaceae, but that genus is retained here in the Dicranaceae.)
1 | Plants robust, with vegetative leaves 3.0–6.5 mm long and not contorted when dry; costa broad, ⅓ to ½ the widest part of the leaf, bordered on both sides by bistratose juxtacostal bands of laminal cells; peristome present; documented only from elevations of >1500 m on the South I. | Holodontium |
1' | Plants small to medium-sized, with vegetative leaves ≤4.0 mm and much contorted when dry; costa narrower, <¼ the widest part of the leaf, not bordered by bistratose laminal cells; peristome present or absent; documented from a wide elevational range on both main islands | 2 |
2 | Peristome absent; capsules immersed to weakly exserted relative to the perichaetial leaves, strongly ribbed when dry; alar cells not differentiated; central strand absent | Amphidium |
2' | Peristome present (but fugacious); capsules exserted on an elongate seta, smooth or wrinkled when dry; alar cells inflated; central strand present | Dicranoweisia |
Category | Number |
---|---|
Indigenous (Non-endemic) | 4 |
Total | 4 |