- ≡ Todea ×intermedia B.S.Williams ex T.Moore., Gard. Chron. 29: 813 (1869)
Leptopteris hymenophylloides and L. superba frequently occur together in wet, forested areas, and they hybridise readily. Plants of Leptopteris ×intermedia are intermediate in morphology between the two parent species (Brownsey 1981, fig. 1). They often have a degree of hybrid vigour and fronds are frequently larger than those of either parent species. The laminae are generally elliptic in outline, like L. superba, but the basal pair of pinnae are longer and the primary pinnae broader than in that species. The ultimate laminal segments are sometimes bent at 90º to the plane of the frond, but not as frequently as in L. superba.
n = 22 (Brownsey 1981). Hybrids show regular bivalent formation at meiosis and have spores of normal appearance, suggesting some degree of fertility (Brownsey 1981).