Classification
 Nomenclature
Scientific Name:
Fissidens tenellus var. australiensis (A.Jaeger) J.E.Beever & I.G.Stone, New Zealand J. Bot. 37: 651 (1999)
Synonymy:
  • Conomitrium perpusillum Müll.Hal. & Hampe in Hampe, Linnaea 28: 214 (1856)
  • Fissidens australiensis A.Jaeger, Enum. Fissident. 24 (1869) nom. nov. pro Conomitrium perpusillum Müll.Hal. & Hampe 1856 (non Fissidens perpusillus Mitt. 1859)
  • Fissidens perpusillus (Müll.Hal. & Hampe) Mitt., Trans. & Proc. Roy. Soc. Victoria 19: 92 (1882) nom. illeg., non Fissidens perpusillus Mitt. 1859
Lectotype: Australia, Victoria, Sealers’ Cove, F. Mueller 113, NY (Designated by Stone 1994a, p.173.) Isolectotypes: BM! MEL!
Etymology:
The varietal epithet, from australis (southern or Australia) + -ensis (inhabiting), is derived from the provenance of this taxon.
 Description

Leaves lanceolate; apex acute; vaginant laminae open; margins denticulate throughout. Costa failing 3–4 cells below leaf apex, bryoides-type in cross-section (although an occasional stereid may be seen on the adaxial face).

Perichaetial leaves with distal region of vaginant laminae margins regularly denticulate, vaginant lamina cells little differentiated. Setae 1.8–3.0 mm. Peristome teeth 24–33 µm wide at base, with ornamentation as per species. Calyptra smooth to slightly scabrous. Spores 10–14 µm.

 Illustrations
 Recognition

Although intermediates with the type variety occur, in its representative form var. australiensis is easily distinguished; it has broader leaf apices with a failing costa, more even serrations on the vaginant lamina margins, smaller vaginant lamina cells, a more or less smooth calyptra, narrower peristome teeth, and larger spores. Distinction at the varietal level is thus appropriate.

 Distribution

K; NI: N Auckland, including offshore islands (PK, HC, LB, GB, RT), S Auckland, Wellington; SI: Nelson (including D’U), Marlborough, Westland, Otago (Catlins); St.

Australasian. Tasmania*, mainland Australia*, New Caledonia*.

 Habitat

Common as an epiphyte or on humic substrates, such as rotting wood, the bases of clumps of Astelia spp., or on decomposing bone; occasionally on soil or rock. It is sometimes found with the only other epiphytic N.Z. Fissidens, F. hyophilus. Additional moss associates include Braithwaitea sulcata, Calyptrochaeta brownii, Dicranoloma menziesii, Haplohymenium pseudotriste, Hymenodon pilifer, Macromitrium spp., and Zygodon intermedius.

Recorded from near sea level to 760 m (Taupō, S Auckland L.D.).

 Biostatus
Indigenous (Non-endemic)
 Bibliography
Beever, J.E. 2014: Fissidentaceae. In: Heenan, P.B.; Breitwieser, I.; Wilton, A.D. (ed.) Flora of New Zealand — Mosses. Fascicle 8. Manaaki Whenua Press, Lincoln.
Beever, J.E.; Malcolm, B.; Malcolm, N. 2002: The Moss Genus Fissidens in New Zealand: an illustrated key. Micro-Optics Press, Nelson.
Beever, J.E.; Stone, I.G. 1999: Studies of Fissidens (Bryophyta: Musci): new taxa and new records for New Zealand. New Zealand Journal of Botany 37: 643–657.
Catcheside, D.G. 1980: Mosses of South Australia. Government Printer, Adelaide.
Hampe, E. 1856: Plantae Muellerianae. Musci frondosi Australasia felici lecti. (Continuatio). Linnaea 28: 203–215.
Jaeger, A. 1869: Enumeratio Generum et Specierum Fissidentacearum adjectis nonnullis adnotationibus de earum litteratura et distributione geographica. Zollikoferi, St. Gallen.
Mitten, W. 1882: Australian mosses, enumerated by William Mitten, Esq. Transactions and Proceedings of the Royal Society of Victoria 19: 49–96.