Classification
 Nomenclature
Scientific Name:
Eurhynchium pulchellum (Hedw.) Jenn., Man. Mosses W. Pennsylvania 350 (1913)
Synonymy:
  • Hypnum pulchellum Hedw., Sp. Musc. Frond. 265 (1801)
Type: Europe. Not seen.
Etymology:
The epithet pulchellum means pretty; it seems a curious name for such a non-descript plant.
 Description

Plants yellow-green, lustrous, forming interwoven mats on soil. Stems to c. 25 mm long, densely and irregularly branched, not terete. Branches mostly short and simple, 5–7(–10) mm, not or weakly complanate in N.Z. material. Stem leaves broadly ovate and acute, narrowed to insertion but not cordate, nearly flat, neither plicate nor decurrent, coarsely serrulate throughout, with a stout costa that lacks a terminal spine, c. 1 mm × 0.5 mm; mid laminal cells narrowly rhombic, firm-walled, smooth, mostly 30–45 × 6–9 μm; cells at apex shorter; alar cells differentiated, forming a medium-sized group, subquadrate to short oblong-rectangular, merging gradually with laminal cells. Branch leaves differentiated, spreading, smaller and more narrowly ovate, ± evenly tapered to an acute apex, with a stout costa only weakly tapering above and ending with a prominent abaxial spine, coarsely serrulate throughout, not or very weakly complanate, mostly 0.8–0.9 × 0.3–0.4 mm; mid laminal cells as per stem leaves, the cells at leaf apex markedly shorter and more rhombic; alar cells moderately differentiated, those at extreme corners ± oblong-rectangular and firm-walled.

Reportedly phyllodioicous. Perichaetia scattered on stems, with perichaetial leaves ecostate and widely spreading. Perigonia not seen in N.Z. material. Setae elongate, smooth. Operculum long-rostrate (only one seen in N.Z. material).

 Distribution

NI: Wellington (Somes I.). Known from a single N.Z. collection. 

Presumably adventive. Widespread in the northern hemisphere.

 Habitat

The sole collection was from track-side soil in coastal scrub of Corynocarpus laevigatus at 30 m elev. The restriction of this rather characterless (at least in the field) species to a single locality in Wellington Harbour seems unlikely, and further collections from other areas might reasonably be expected.

 Biostatus
Exotic
 Notes

The branch leaf alar cells here are more elongate and oblong than those of the northern hemisphere herbarium specimens available for comparison, but compare well to Buck’s (1998) illustrations. The species is highly variable in Europe, and numerous variants have been proposed; no attempt has been made to assign the N.Z. collection to any variants.

 Bibliography
Buck, W.R. 1998: Pleurocarpous mosses of the West Indies. Memoirs of the New York Botanical Garden 82: 1–400.
Crum, H.A.; Anderson, L.E. 1981: Mosses of Eastern North America. Columbia University Press, New York.
Fife, A.J. 2020: Brachytheciaceae. In: Smissen, R.; Wilton, A.D. (ed.) Flora of New Zealand – Mosses. Fascicle 46. Manaaki Whenua Press, Lincoln.
Hedwig, J. 1801: Species Muscorum Frondosorum descriptae et tabulis aeneis lxxvii coloratis illustratae. Barth, Leipzig.
Jennings, O.E. 1913: A Manual of the Mosses of Western Pennsylvania. Pittsburgh.